The study of phytocenoses of transitional environmental conditions in the Pre-Baikal Region enabled to reveal modern tendencies of vegetation development in different Pre-Baikal sites. Such cenoses can serve as a regional model for indication of existing processes and of changes in vegetation structure which occurred under different environmental conditions. It is found out that ecotones and cenoses reflecting the paragenese in vegetat ion structure indicate structural-dynamic peculiarities of the organization of Pre-Baikalian vegetation cover as a whole. They are valid for classification and characterize intrazonal differences in vegetation structure at vaste territories. Ecotones and paragenese in the vegetation structure in the studied area reflect physical and geographical conditions of their formation for a definite time period.
The problem of assessment of ecosystems state and of the forecast of their changes in a whole and in the structure of vegetation covet in particular is a base of modern biogeographical, geobotanical, biogeocenotic and ecological studies. The main task is to study spatial-temporal self-organization of phytocenoses reflecting practically all changes in the environment during a concrete time period. To resolve it, a synthesis of different ways of their studies is necessary rather than a detailed analysis of separate components and elements of natural ecosystems. With such an approach, a phytocenosis is considered as a system, which forms and develops as a complex unit and forms mutually initiated links of matter and energy exchange with systems of other hierarchical level and ecotopes conditions both for separate species and for cenoses. The methods of determination of long-term trends of dynamics of vegetation cover structure are at different stages of development causing to researchers a series of problems―from choice of conceptual basis to notions and terms establishment for the revealed processes and states occurring in the vegetation cover and in whole ecosystems. The resolution of such tasks makes obviously necessary to correct the existing understanding of processes occurring in the vegetation cover and allows to find out a vector of their development within the system of natural factors at any territory. This results in appearing of basis to establish the age, the place and the role of current states of phytocenoses in successional systems. It is necessary for it to reveal the peculiarities of composition and assortment of phytocenoses resulting from natural environmental dynamics and from anthropogenic impact, especially for the vegetation from contrast environmental conditions.
At present, there is no unique approach to determine a sense of the notion “ecotone” while characterizing the structure of the vegetation cover under the condition of contrast (transitional) environmental conditions. This is partly due to the insufficiency of criteria determining the range of this notion. Due to this fact, this term is used for characterization of a transitional state both of vegetation in a whole environmental zone [
There is a greater misunderstanding for the term “paragenese” while characterizing spatial-temporal organization of vegetation. If in geography, geology, geomorphology and geochemistry it is rather established [
The aim of this work is to find out the peculiarities of spatial-temporal organization of phytocenoses as ecotones between zonal forest-steppe and zonal steppe as well as one of phytocenoses reflecting the paragenes in the vegetation structure in the Baikal Region presenting concrete key sites. The following tasks were being resolved: to find out areological and ecotypological composition of the flora in the cenoses forming under the conditions of steppes extrazonality and at the boundary of forest-steppe and steppe environmental zones; to typify phytocenoses at environments contact sites (transitional natural conditions) between environmental zones, as well as phytocenoses of intrazonal environment differences; to determine vectors of formation of such phytocenoses under modern environmental conditions in the region.
The studied areas (key sites) are the following: middle part of the Selenga River basin (South-Western Trans- Baikal), middle part of the Barguzin Depression (North-Eastern coast of Lake Baikal). We have used in our work the method of field geobotanical survey. To reveal the links of phytocenoses with edaphic conditions in the habitats, we performed a combined soil-geobotanical profiling for the key sites at the studied territory. While classifying vegetation, we used a dominant approach with characterization of horizons species composition, which allows to reveal a spatial structure of phytocenoses from Arctic tundras to arid deserts under the conditions of zonality and height-belts. This approach is natural and fruitful in geobotanical mapping for revealing of the specifics of vegetation cover structure at different levels of its organization. The cenoses were united into associations by structure, dynamics and location similarity. The variable states of these associations were united in turn into epitaxa [
Natural and climatic conditions of the region are presented in numerous specialized maps, atlases and papers and reflect these or those characteristics of regional environments and structural peculiarities of the vegetation cover on the territory of the Baikal Region from first papers by M.F. Korotky [
We believe that it is possible to present only several rather specific characteristics of the factors of phytocenoses formation in the description of environmental conditions directly for each key site which are representative for concrete areas of the whole Baikal region. In the system of geographical zonality by heat and humidity ratio, the latitudinal zonality is primary, and height belts are secondary. Due to changes in solar radiation at the boundary of transition of atmospheric moistening (from oceans deepward to the continent), the latitudinal zonality in mountain systems is substituted by height belts, which is in a definite belt of solar radiation. I.e., in the mountains, the height belts are determined by the intensity of solar radiation of a definite radiation belt on the general background of heat and humidity ratio in the aspect of latitudinal zonality [
The basis vegetation of this key site consists of taiga (boreal) vegetation of Ural-Siberian fratria of formations of mountain taiga pine and larch pine grass-brush forests in combination with pine grass-brush stepped forests and small-bunchgrass steppes of South Siberia formations of forb-cereal and cereal steppes [
We performed an arealogical analysis (see
Geoelemental and ecotypological composition of the basic plant species of the key area | |||||
---|---|---|---|---|---|
Plant species | Geoelement | Ecotype | Plant species | Geoelement | Ecotype |
Pinus sylvestris | ЕА | XM | Dracocephalum ruyschiana | ЕА | XM |
Larix sibirica | ES | MT | Equisetum sylvaticum | CP (HA) | MT |
Betula pendula | ES | MT | Sanguisorba officinalis | CP (HA) | MT |
Spiraea media | ЕА | MT | Poa botryoides | CA | MX |
Cotoneaster melanocarpus | NA | XM | Artemisia absinthum | ЕА | MX |
Rosa acicularis | CP (HA) | MT | Astragalus subfrutocosus | SC | MX |
Carex pediformis | ЕА | MX | Pedicularis venusta | SC | MT |
Carex macroura | NA | MT | Vicia baikalensis | E-A | MT |
Pulsatilla multifida | CP (HA) | MT | Carex duriuscula | АА | XT |
Bupleurum sibiricum | MD | XM | Trifolium pratense | ES | MT |
Scorzonera radiata | CA | XM | Myosotis arvensis | CP (HA) | HM |
Crepis sibirica | EA | MT | Papaver popovii | EN | XM |
Thalictrum foetidum | EA | XM | Taraxacum officinale | SC | MT |
Phlomis tuberosa | ЕА | XM | Allium tenuissimum | SC | XM |
Potentilla bifurca | ЕC | XM | Poa pratensis | ЕА | MT |
Myosotis imitata | CP (HA) | MT | Rumex acetosella | CP (HA) | MT |
Vicia cracca | C-A | MT | Festuca lenensis | NEA | CrX |
Polygala sibirica | ЕА | MT | Artemisia frigida | CP (HA) | CrX |
Draba nemorosa | CP (HA) | MT | Potentilla acaulis | CA | XT |
Galium verum | CP (HA) | MT | Orostachis spinosa | CA | CrX |
Astragalus austrosibiricus | NA | MX | Veronica incana | NA | MX |
Polygonatum odoratum | EА | MT | Aster alpinus | CA | CrX |
Ranunculus borealis | ЕА | MT | Clausia aprica | ЕА | XT |
Fragaria viridis | ES | MT | Potentilla tanacetifolia | SC | XM |
Alopecurus pratensis | ES | MT | Thymus serpyllum | SC | XT |
Trollius asiaticus | SC | MT | Goniolimon speciosum | ЕА | XT |
Filipendula ulmaria | ЕА | MT | Eremogone meyerii | SC | MX |
Urtica dioica | OKh | MT | Heteropappus altaicus | C-A | XM |
Maianthemum bifolium | ЕА | MT | Artemisia laciniata | NA | MX |
Plantago major | CP (HA) | MT | Iris humilis | ЕА | MX |
Stellaria cherleriae | SC | MT | Androsace lactiflora | NA | XM |
Thalictrum foetidum | ЕА | MT | Stipa sibirica | NA | XM |
Trifolium repens | ЕА | MT | Chamaerhodos erecta | NA | MX |
Pyrola rotundifolia | CP (HA) | MT | Trifolium lupinaster | ЕА | MT |
Koeleria cristata | CP (HA) | XT | Alyssum obovatum | CA | CrX |
---|---|---|---|---|---|
Capsella bursa-pastoris | ЕА | MT | Thermopsis lanceolata | ЕА | ГалXM |
Artemisia vulgaris | CP (HA) | MT | Erysimum flavum | NA | MX |
Agrostis gigantea | CP (HA) | MT | Taraxacum mongolicum | E-A | XT |
Atriplex sibirica | C-A | MX | Carex korshinskyi | E-A | MX |
Chelidonium majus | ЕА | MT | Androsace incana | SC | CrX |
Euphorbia discolor | NA | XM | Schizonepeta multifida | NA | MX |
Leymus chinense | C-A | MT | Astragalus adsurgens | E-A | XM |
Calamagrostis epigeios | ЕА | XM | Serratula centauroides | SC | XM |
Plantago media | ЕА | MT | Ptilotrichum tenuifolium | SC | XT |
Stipa krylovii | C-A | MT | Agropyron cristatum | CA | MX |
conditions where one can observe processes of mutual invasion of plants from mountain forest-steppe zone into a steppe zone, and the key site (middle part of the Selenga River basin) is situated within the territory of the contact of Orkhon-Lower Selenga mountain-forest-steppe subprovince and mid-Khalkhass steppe subprovince of Central-Asian (Dauria-Mongolia) subregion of Eurasian steppe area [
In some locations of the cenoses (on the base of geobotanical descriptions) mosses of a wide ecological amplitude characteristic for polydominant light coniferous forests occur extremely rarely, synusially. These are such species as Abietinella abietina (Turn.) Fleisch., Polytrichum piliferum Hedw., Ceratodon purpureus (Hedw.) Brid., Funaria hygrometrica Hedw., Pylaisia polyantha (Hedw.) BSG., Orthotrichum rupestre Schleich. ex Schwaeger. Essentially, these cenoses form an interzonal ecotone. Consequently, they can be used for monitoring of spatial variability of vegetation cover in environmental zones of South-Western Trans-Baikal on the background of climate dynamics in the region during last decades.
The structure of forb stepped forests in the studied area reflects the tendences of increase of undergrowth amount since 1970ies. The dynamics of reconstitution processes in the forests under the conditions of zonal forest-steppe reflects differentiation of surface cover by horizons, abundant undergrowth with tendency to further forestation of open steppe sites both within the zone of mountain forest-steppes and at contact site with the steppe zone. The performed soil-geobotanical profiling allowed to reveal a rather tight correlation of vegetation structure with soils. Forests and steppe cenoses are rather closely linked with soils types characteristic for definite elements of the relief structure; this is appropriate to mountain forest-steppes where vertical belts are manifested (or height zonality, by: Reimers [
If we consider present-day forestation processes of steppe territories within a zonal forest-steppe (the Selenga R. basin) at a contact site with Central Asia steppe zone, this will be probably mainly spatial re-distribution of territories occupied by forest and steppe cenoses with time with possible change of the forest-steppe boundary in latitudinal direction due to climatic fluctuations for a concrete time period on the background of forcing of processes of anthropogenic impact onto the environment. Under the conditions of contact between zonal steppe and zonal forest-steppe, phytocenoses which are just to be called ecotones form―these are transitional cenoses between environmental zones. An interzonal ecotone, in this case―one between zonal steppes and zonal forest- steppes, reflects a spatial-temporal variability of vegetation structure at definite environmental and climatic conditions. At humidity increase, a shift of forest-steppe boundary towards the steppes zone (in latitudinal direction) is observed as forestation of steppe territories both within forest-steppe zones and at a contact site of steppe and forest-steppe zones, respectively. Here one can suppose that at income of a dry period, one forecast extention of steppes zone toward the forest-steppe zone with extention of steppe territories within forest-steppes. It occurred just in recent past―at different Holocene stages in Siberia [
There is a following conclusion for this key site. Due to climate change (increase of mean annual temperatures and humidity increase at last stages of vegetation), on the background of the past period of aridization (Late Holocene), processes of forests reconstitution instead of steppe territories occur everywhere. On the background of pasturage charge decrease during last decades, the grasses are reconstituted rather slowly (division into horizons, increase of projective cover, increase of species diversity). Due to these processes, change of forest-steppes boundary in the latitudinal direction В (forestation of steppe territories everywhere) is possible; this reflects a permanent state of “steppe islands” for a concrete time period. This is confirmed as well by the availability of sod-forest (forest zone soils) soils with processes of soils formation according to the “forest” type. It is to consider here increase of anthropogenic impact, mainly pasturage regime accompanied often by periodic fires (grassland fires) as a constraint factor for vegetation formation.
Characteristic forests for the Barguzin Depression are mountain taiga ones of Baikal-Dzhugdzhur formation of Argada fratria together with piedmont-depressional light coniferous ones of South-Siberian formations. Larix- pine suffruticous-green mosses forests with underbrush of rhododendron and Duschekia fruticosa form along the depression boards and slopes aprons of mountain surrounding (Barguzin and Argada Ridges). The bottom of Barguzin Depression itself is mainly ploughed up, and at yields margin, forbs-cereals cenoses of South-Siberian formations occur [
Probably, in this case we have to speak about climatogenic succession as about a process where there is a given reference point of phytocenoses state under concrete physical-geographical conditions for a definite territory for a concrete time period leading to formation of a qualitatively different morphology of a cenoses differing from one available at the reference point. From our viewpoint, the climatogenic successions reflect paragenese in vegetation organization, which is suggested by the results of the studies performed, including the analysis of the composition of geoelements and ecotypes of plants species in the cenoses from the studied area. Main feature of vegetation structure in Barguin Depression in the whole is formation of pine forests together with steppe cenoses of depression bottoms and of slopes aprons of different expositions. Steppe cenoses consist of large cereals steppes with domination of Stipa krylovii Roshev., Agropyron cristatum (L.) Beauv. The sites occupied by these cenoses were ploughed up in the middle period of the last century. It is noticed that here there are no fescue (Festuca lenensis Drobov) steppes characteristic for the whole Trans-Baikal, but there are instead of them well developed steppes with domination of Artemisia obtusiloba ssp. subviscosa (Turcz. ex Besser) Krasnov and with large distribution of Artemisia frigida Willd. The cereal component of the cenoses includes such species as Agropyron cristatum (L.) Beauv., Poa botryoides (Trin. ex Griseb.) Kom., Cleistogenes squarrosa (Trin.) Keng. Among forbs, there are in abundance Potentilla acaulis L., Oxytropis oxyphylla (Pallas) DC., Papaver nudicaule L., Potentilla bifurca L., Chamaerhodos erecta (L.) Bunge., Delphinium grandiflorum L. and Thymus serpyllum L. There are well developed cenoses with domination of Poa botryoides (Trin. ex Griseb.) Kom., Bupleurum bicaule Helm. and Artemisia obtusiloba ssp. subviscosa (Turcz. ex Besser) Krasnov. There are significant groups with domination of Stipa krylovii Roshev. A significant cenoses differentiation by their species and quantitative composition related to concrete depression territories is observed in the spatial structure of Barguzin Depression.
The base of modern vegetation of this key site is larch (Larix sibirica Ledeb.)-pine (Pinus sylvestris L.) forbs-stepped thinned forests with pine undergrowth and sprouts in combination with steppe cenoses in which there are clumps of pine undergrowth and sprouts, especially on the territories not impacted by ploughing up. The main horizon is dominated by pine, larch is co-dominant. The undergrowth is abundant and is spread out of the canopy. The brush horizon consists of Rhododendron dahuricum L., Spiraea media Franz Schmidt., Rosa acicularis Lindley. The surface cover is characterized by Dianthus versicolor Fischer ex Link, Veronica incana L., Allium tenuissimum L., Stipa krylovii Roshev., Artemisia obtusiloba ssp. subviscosa (Turcz. ex Besser) Krasnov, Galium verum L., Patrinia rupestris (Pallas) Dufr., Elytrigia repens (L.) Nevski. There are synusias of Abietinella abietina (Turn.) Fleisch., Rhytidium rugosum (Hedw.) Kindb. Typical for polydominant dark coniferous―light coniferous Pre-Baikalian taiga. Basic modern vegetation of this key sites are forbs-stepped thinned pine forests with pine undergrowth in combination with steppe cenoses with clumps of pine undergrowth and sprouts, especially on the territories not impacted by ploughing up. Modern tendencies are characterized by forestation processes everywhere in steppe space. Under the conditions of contact of forests and extrazonal steppes of depression types, cenoses with plants species from different environmental confinedeness form resulting from environmental specifics.
The dominant families (Asteraceae, Poaceae, Cyperaceae, Fabaceae, Apiaceae, Brassicaceae, Ranunculaceae, Caryophyllaceae) form major part of plants species mentioned in geobotanical descriptions of different years and vegetation periods for the key site. This set of dominant families is very characteristic for boreal floras and is similar to the family spectrum for East Siberian flora [
Among main (the most occurring plants species in geobotanical descriptions) floristic composition (see
Essentially, the cenoses in the studied area reflect the peculiarities of relations and spatial variation between forests and extrazonal steppe cenoses. These cenoses formed by plants species with different ecological amplitudes respond very rapidly and reflect visually quite distinctly changes in ecotopes conditions at topological and regional levels of environment organization. Such cenoses can serve as representative models reflecting structural-dynamical peculiarities of the vegetation in the region on the background of climate dynamics during last decades. The comparative analysis of areas occupied by forest and steppe cenoses showed tendencies to a gradual forestation of steppe space, especially in lower parts of graded ridge slopes in the depression surroundings. Steppe cenoses (listed on the base of summarized geobotanical descriptions and vegetation periods) are forbs-cereal plants groups with inclusions in them of pine of different age (5 - 25 y.o.) heterogenous by distribution territory. Forests (listed on the base of summarized geobotanical descriptions and vegetation periods) are larch-pine, pine brush forbs forests together with open stepped ones with pine undergrowth forests at the boundary with extrazonal (depression type) steppes of bottom and slopes aprons on the depression boards. Ploughed fields are soils ploughed up in 1950ies, representing now perennial yields, territories with tendencies of formation here of plants groups of steppe types at first stage. Pine undergrowth is rare.
Modern tendencies of vegetation development in the studied area reflect a transitional character of steppes with tendencies with tailing of boundary between forest and steppe cenoses. The appearing of pine undergrowth in extrazonal steppes can suggest that in the region, together with forests reconstitution dynamics, the structure of steppe cenoses changes as a response to changing climatic situation. This is just confirmed by natural forestation of extrazonal steppes in the whole Barguzin Depression. Further cenoses development in the studied area from the viewpoint of structural changes on the background of climatic factors fluctuations will go towards forests formation with presence of small territories occupied with xerophyte-petrophyte grass groups on stony substrates and ridges. These processes may be constrained by the character of anthropogenic impact or by abrupt climate changes in the region making climate dryer.
Forestation of extrazonal steppe of Barguzin Depression (North-Eastern coast of Lake Baikal ) as paragenesis (object) suggests changes in natural climatic conditions in the region, when steppe territories in some way “compress” by their territory for a concrete time period. It means that for last decades, this process has a regional scale. And in this case, as it was mentioned before, it is to notice that “A forest of this or that kind, composition or structure will obviously cover earlier or later steppe spaces at least by park plants but with keeping of small steppe “islands” due to increase of climate dryness in the beginning of the century” [
The cenoses at such key site such as middle part of the Selenga River basin, South-Western Pre-Baikal―territory on the junction site of mountain forest-steppes and steppe zone of Central Asian (Daurian-Mongolian) subregion of a steppe region―represent an interzonal ecotone. This is suggested by the results of analysis of geoelemental and ecotypological (ecotypes) compositions, of belt-zonal groups of plants species and the composition of dominant families. By species composition, key site cenoses are more related to the forest-steppe zone but with presence of species representing North-Asian steppes. For the territory of the studied area, zonal forest-steppe soils are characteristic. Climate change during last decades in this region is manifested by humidity and temperature increase. Decrease of anthropogenic pression results as well both in forestation of steppe space within the
Geoelemental and ecotypological composition of the basic plant species of the key area | |||||
---|---|---|---|---|---|
Plant species | Geoelement | Ecotype | Plant species | Geoelement | Ecotype |
Pinus sylvestris | ЕА | XM | Artemisia frigida | CP (HA) | CrX |
Populus tremula | ЕА | XM | Thymus serpyllum | SC | XT |
Rhododendron dahuricum | MD | MT | Potentilla acaulis | CA | XT |
Spiraea media | ЕА | MT | Artemisia gmelinii | NA | CrX |
Artemisia subviscosa | EN | MX | Bupleurum scorzonerifolium | E-A | XM |
Dianthus versicolor | ЕА | MX | Leontopodium ochroleucum | C-A | CrX |
Veronica incana | NA | MX | Phleum phleoides | ES | XM |
Orostachys spinosa | CA | CrX | Spiraea flexuosa | MD | MT |
Iris ruthenica | NA | MT | Agropyron cristatum | CA | XM |
Poa botryoides | NA | MX | Artemisia commutata | NA | MX |
Allium tenuissimum | SC | XM | Scabiosa comosa | SC | XM |
Rosa acicularis | CP (HA) | MT | Linum sibiricum | ЕА | MX |
Bergenia crassifolia | SC | MT | Scorzonera radiata | NA | XM |
Vaccinium vitis-idaea | CP (HA) | XM | Draba nemorosa | CP (HA) | MT |
Trollius asiaticus | SC | MT | Poa sibirica | NA | XM |
Chelidonium majus | ЕА | MT | Bupleurum bicaule | MD | CrX |
Vicia baicalensis | E-A | MT | Galium boreale | CP (HA) | MT |
Sanguisorba officinalis | CP (HA) | MT | Myosotis suaveolens | ЕА | MX |
Leucanthemum vulgare | ЕА | XM | Leontopodoium leontopoides | MD | CrX |
Goniolimon speciosum | ЕА | HalXM | Koeleria cristata | CP (HA) | XT |
Stipa krylovii | C-A | XM | Schizonepeda multifida | NA | MX |
Cotoneaster melanocarpus | ЕА | XM | Astragalus versicolor | SC | XM |
Carex macroura | NA | MT | Androsace incana | SC | CrX |
Carex pediformis | ЕА | MX | Galium verum | CP (HA) | MT |
Polygonatum sibiricum | CA | MT | Thalictrum foetidum | ЕА | XM |
Patrinia rupestris | SC | CrX | Pulsatilla multifida | CP (HA) | XM |
Phlomis tuberosa | ЕА | XM | |||
Aster alpinus | CA | CrX | |||
Artemisia laciniata | NA | MX | |||
Carex korshinskyi | E-A | MX |
steppe zone and in extention of trees species towards steppes. Thus, there is a tendency of forest-steppe zone shifting in latitudinal direction with formation to some degree of light coniferous forests of zonal type. At the boundaries of environmental zones where ecotones form as transitional systems at changing heat and humidity ratio, their considerable territorial “widening” (or “narrowing” at other scenarios of climate changes) occurs. This is just seen at the sites of relative spatial “widening” of the ecotone between forest-steppe (mountain forest-steppes) and steppe environmental zones of South-Eastern Trans-Baikal.
Species composition and structure of phytocenoses at the key site (within Barguzin Depression) reflect the specifics of vegetation formation under the conditions of extrazonality of steppe cenoses contacting with polydominant light coniferous forests, often with inclusions of dark coniferous species in the undergrowth and in the second horizon. There are among forests vegetation groups where species composition is based on xerophytes of steppe environmental zone. Soils of the key site (Barguzin Depression) are characterized as extrazonal (cryoarid) ones; they do not reflect direct connection with species composition and phytocenoses types on this territory. Both forest and steppe cenoses develop on the same soils; it was revealed at combined soil-geobotanic profiling of key sites. Such cenoses reflect paragenese (object) in organization of vegetation cover on a concrete territory. Paragenese in vegetation structure means cenoses reflecting structure and dynamics of environmental conditions of regional-topological level of environment organization within environmental zones during a definite time period on a concrete territory. Decrease of areas occupied by extrazonal steppe cenoses in forest zone during last decades allows to speak about rather considerable climate changes and about anthropogenic impact in the region. Forest cenoses with rather sustainable renewal, often with synusia of mosses characteristic for polydominant dark coniferous―light coniferous taiga form intensively among steppe sites. Under the conditions of steppes extrazonality and of non-expressed mountain steppe and mountain-forest-steppe belts, particular cenoses which are to be called taiga-steppe ones form in the studied area. Taking into account the peculiarities of modern state and development vector of taiga-steppe cenoses of the studied area (Barguzin Depression), we have to wait for formation of light coniferous taiga with dark coniferous species in perspective, as there are here tendencies to strengthening of their positions such as considerable amount of dark coniferous species in the second horizon and in pine forests undergrowth on the studied territory. Under paragenese conditions (within environmental zones) as in definite environments differing by heat and humidity ratio, by radiation balance at climate change, mainly towards precipitation increase, an environmental site is “narrowing” (in our case this is forestation of extrazonal steppes within zonal taiga) during a concrete time period at a definite site. This was just found out at the example of vegetation structure of a key site in the Baikal Region―Baikal Depression. This environmental site can “widen” as well in space due to increase of xerophytation processes (at increase of climate dryness) within taiga zone vegetation cover. However, these processes will depend on vector and character of anthropogenic impact on the background of climate dynamics, mainly on moistening.
The ecological and biocenological role of ecotones and cenoses reflecting paragenese (object) in the vegetation structure of the Baikal region is in revealing of phytocenotic and typological diversity of cenoses, in indication of structural-dynamic organization and in forecast of vegetation development under concrete physical geographical conditions at vaste territories including zonal and intrazonal differences of environments determining different degree of natural and anthropogenic resistance of the whole vegetation cover.
Alexander Sizykh, (2016) Arealogical and Ecotypological Compositions of the Flora of Plant Communities at Environment Contact Sites (Based on Some Pre-Baikal Sites). Open Access Library Journal,03,1-12. doi: 10.4236/oalib.1102823